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Biodiversity change through geological time

The number of species on the planet, or in any geographical area, is the result of an equilibrium of two evolutionary processes that are ongoing: speciation and extinction. Both are natural “birth” and “death” processes of macroevolution. When speciation rates begin to outstrip extinction rates, the number of species will increase; likewise, the number of species will decrease when extinction rates begin to overtake speciation rates. Throughout Earth’s history, these two processes have fluctuated—sometimes leading to dramatic changes in the number of species on Earth as reflected in the fossil record ( [link] ).

The graph plots percent extinction occurrences versus time in millions of years before present time, starting 550 million years ago. Extinction occurrences increase and decrease in a cyclical manner. At the lowest points on the cycle, extinction occurrences were between 2% and 5% percent. Spikes in the number of extinctions occurred at the end of geological periods: end-Ordovician, 450 million years ago; end-Devonian, 374 million years ago; end-Permian, 252 million years ago; end-Triassic, 200 million years ago; and end-Cretaceous, 65 million years ago. During these spikes, extinction occurrences approximately ranged from 22% to 50%.
Percent extinction occurrences as reflected in the fossil record have fluctuated throughout Earth’s history. Sudden and dramatic losses of biodiversity, called mass extinctions, have occurred five times.

Paleontologists have identified five strata in the fossil record that appear to show sudden and dramatic (greater than half of all extant species disappearing from the fossil record) losses in biodiversity. These are called mass extinctions. There are many lesser, yet still dramatic, extinction events, but the five mass extinctions have attracted the most research. An argument can be made that the five mass extinctions are only the five most extreme events in a continuous series of large extinction events throughout the Phanerozoic (since 542 million years ago). In most cases, the hypothesized causes are still controversial; however, the most recent event seems clear.

The five mass extinctions

The fossil record of the mass extinctions was the basis for defining periods of geological history, so they typically occur at the transition point between geological periods. The transition in fossils from one period to another reflects the dramatic loss of species and the gradual origin of new species. These transitions can be seen in the rock strata. [link] provides data on the five mass extinctions.

This table shows the names and dates for the five mass extinctions in Earth’s history.
Mass Extinctions
Geological Period Mass Extinction Name Time (millions of years ago)
Ordovician–Silurian end-Ordovician O–S 450–440
Late Devonian end-Devonian 375–360
Permian–Triassic end-Permian 251
Triassic–Jurassic end-Triassic 205
Cretaceous–Paleogene end-Cretaceous K–Pg (K–T) 65.5

Explore this interactive website about mass extinctions.

The pleistocene extinction

The Pleistocene Extinction is one of the lesser extinctions, and a recent one. It is well known that the North American, and to some degree Eurasian, megafauna    , or large animals, disappeared toward the end of the last glaciation period. The extinction appears to have happened in a relatively restricted time period of 10,000–12,000 years ago. In North America, the losses were quite dramatic and included the woolly mammoths (last dated about 4,000 years ago in an isolated population), mastodon, giant beavers, giant ground sloths, saber-toothed cats, and the North American camel, just to name a few. The possibility that the rapid extinction of these large animals was caused by over-hunting was first suggested in the 1900s. Research into this hypothesis continues today. It seems likely that over-hunting caused many pre-written history extinctions in many regions of the world.

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Source:  OpenStax, Bi 101 for lbcc ilearn campus. OpenStax CNX. Nov 28, 2013 Download for free at http://legacy.cnx.org/content/col11593/1.1
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