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A micrograph of an oval cell with many short projections.
This specimen of the ciliate Balantidium coli is a trophozoite form isolated from the gut of a primate. B. coli is the only ciliate capable of parasitizing humans. (credit: modification of work by Kouassi RYW, McGraw SW, Yao PK, Abou-Bacar A, Brunet J, Pesson B, Bonfoh B, N’goran EK&Candolfi E)
Paramecium cell with short strands on the outside labeled cilia. An indent in the outer layer is labeled cytostome. The outside edge of the cytostome is an indent in the cell labeled oral groove. A sphere inside the cell at the base of the cytostome is labeled food vacuole, another nearby sphere is labeled cytoproct. A smaller opening in the cell is labeled anal pore. A star shaped structure inside the cell is labeled contractile vacuole. A large oval is labeled macronucluus and a smaller oval is labeled micronucleus.
Paramecium has a primitive mouth (called an oral groove) to ingest food, and an anal pore to excrete it. Contractile vacuoles allow the organism to excrete excess water. Cilia enable the organism to move.
A micrograph of long trumpet shaped cells. The wide part of the cell has an oval structure labeled cytostome and small projections labeled cilia.
This differential interference contrast micrograph (magnification: ×65) of Stentor roeselie shows cilia present on the margins of the structure surrounding the cytostome; the cilia move food particles. (credit: modification of work by “picturepest”/Flickr)

Öomycetes have similarities to fungi and were once classified with them. They are also called water molds . However, they differ from fungi in several important ways. Öomycetes have cell walls of cellulose (unlike the chitinous cell walls of fungi) and they are generally diploid, whereas the dominant life forms of fungi are typically haploid. Phytophthora , the plant pathogen found in the soil that caused the Irish potato famine , is classified within this group ( [link] ).

A photograph of an insect covered in white fuzz labeled water mold.
A saprobic oomycete, or water mold, engulfs a dead insect. (credit: modification of work by Thomas Bresson)

Excavata

The third and final supergroup to be considered in this section is the Excavata, which includes primitive eukaryotes and many parasites with limited metabolic abilities. These organisms have complex cell shapes and structures, often including a depression on the surface of the cell called an excavate. The group Excavata includes the subgroups Fornicata , Parabasalia , and Euglenozoa . The Fornicata lack mitochondria but have flagella. This group includes Giardia lamblia (also known as G. intestinalis or G. duodenalis) , a widespread pathogen that causes diarrheal illness and can be spread through cysts from feces that contaminate water supplies ( [link] ). Parabasalia are frequent animal endosymbionts; they live in the guts of animals like termites and cockroaches. They have basal bodies and modified mitochondria (kinetoplastids). They also have a large, complex cell structure with an undulating membrane and often have many flagella. The trichomonads (a subgroup of the Parabasalia) include pathogens such as Trichomonas vaginalis , which causes the human sexually transmitted disease trichomoniasis . Trichomoniasis often does not cause symptoms in men, but men are able to transmit the infection. In women, it causes vaginal discomfort and discharge and may cause complications in pregnancy if left untreated.

The Euglenozoa are common in the environment and include photosynthetic and nonphotosynthetic species. Members of the genus Euglena are typically not pathogenic. Their cells have two flagella, a pellicle , a stigma (eyespot) to sense light, and chloroplasts for photosynthesis ( [link] ). The pellicle of Euglena is made of a series of protein bands surrounding the cell; it supports the cell membrane and gives the cell shape.

Questions & Answers

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Source:  OpenStax, Microbiology. OpenStax CNX. Nov 01, 2016 Download for free at http://cnx.org/content/col12087/1.4
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