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Schooling in fish  (Page 2/10)

Species used to study of schooling
Species Habitat Notes Researcher(s) Cited In Paper
Spottail Shiner ( Notropis hudsonius ) Freshwater Rivers A migratory and strongly schooling fish Dr. Benoni Seghers,1981
Norwegian Herring ( Clupea harengus L. ) Northeast Atlantic One of few very inflexible fish in terms of schooling behavior Dr. TJ Pitcher, 1991
Trinidadian Guppy ( Poecilia reticulata ) Freshwater streams Small fish that is easily manipulated and therefore good for experimental use Dr. Anne Magurran, 1991, 1994.
Three spined stickleback ( Gasterosreus aculeaius ) Freshwater lakes Small fish that is common to North America and Europe Dr. V Kaitala, Dr. E Ranta, 2006
Goldfish ( Carassius auratus ) Freshwater Have the ability to school, but rarely do so Dr. Anne Magurran, 1982
Minnows ( Phoxinus phoxinus ) Freshwater Have the ability to school, but rarely do so Dr. Anne Magurran, 1982
Juvenile roach ( Rutilus Rutilus ) Freshwater Does not always school, prefers shallow water Dr. Dirk Bumann, 2004
Northern bluefin tuna ( Thunnus thynnus ) Atlantic Ocean Fast swimmers who often school Dr. TJ Pitcher, 1999
North esk salmon ( Salmo salar ) North Esk Freshwater River Migrating fish that are a prime target of fishermen Dr. ADF Johnstone,1995
Eastern mosquitofish ( Gambusia holbrooki ) Freshwater Famous for sexual schooling preferences Dr. Angelo Bizazza, 2007
Banded killifish ( Fundulus Diaphanus ) Freshwater Adjust schooling behavior to resource availability often Dr. DJ Hoare, 2004
Juvenile chum salmon( Oncorhynchus keta ) Freshwater rivers Usually always school Dr. Bori Olla and Dr. Clifford Ryer, 1991
French grunts ( Haemulon flavolineatum ) Coral Reefs Famous for involvement in mixed schooling Dr. E Ranta, 1994
Golden shiners ( Notemigonus crysoleucas) Freshwater lakes Used in communication studies often Dr. E Ranta, 1994

Schooling was initially thought to be a behavior with little structure or adaptive significance (Keenleyside 1955), however further study has revealed an intricate and developed structure behind the school. Individuals are capable of plastic behavior in terms of when and where they school. The banded killifish ( Fundulus Diaphanus ), which live in isolated populations, stay in close proximity of one another and when there is a shortage of food or a predation threat, they quickly band together and school. Staying close together affords them this flexibility and is seen as a function of external stimuli (Hoare et al 2004). Scientists have discovered that schools are much more complex in their structure than originally thought, lacking almost any randomness. Instead, individuals compete for positions within the school, with edge positions typically falling to those with the least fitness (Hamilton 1970). These schools are often are controlled by signals within the group from neighbor to neighbor. The signals are used to direct traffic, indicate the presence of food, aid recognition of school members, and send out alarm calls in the face of predators (Magurran 1994). Consequently, the reactions to finding food or encountering a predator are well orchestrated and coordinated. Fish may or may not school, and these choices depend on food availability, predator density, and sometimes for females, even the level of sexual harassment present. Here we examine the evolutionary reasons for schools, how these schools are able to adapt to environmental changes, and the known aspects of school mechanics .

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OpenStax, Mockingbird tales: readings in animal behavior. OpenStax CNX. Jan 12, 2011 Download for free at http://cnx.org/content/col11211/1.5
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