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Photo is of a snapdragon with a pink flower.
These pink flowers of a heterozygote snapdragon result from incomplete dominance. (credit: “storebukkebruse”/Flickr)

Codominance

A variation on incomplete dominance is codominance    , in which both alleles for the same characteristic are simultaneously expressed in the heterozygote. An example of codominance is the MN blood groups of humans. The M and N alleles are expressed in the form of an M or N antigen present on the surface of red blood cells. Homozygotes ( L M L M and L N L N ) express either the M or the N allele, and heterozygotes ( L M L N ) express both alleles equally. In a self-cross between heterozygotes expressing a codominant trait, the three possible offspring genotypes are phenotypically distinct. However, the 1:2:1 genotypic ratio characteristic of a Mendelian monohybrid cross still applies.

Multiple alleles

Mendel implied that only two alleles, one dominant and one recessive, could exist for a given gene. We now know that this is an oversimplification. Although individual humans (and all diploid organisms) can only have two alleles for a given gene, multiple alleles may exist at the population level such that many combinations of two alleles are observed. Note that when many alleles exist for the same gene, the convention is to denote the most common phenotype or genotype among wild animals as the wild type (often abbreviated “+”); this is considered the standard or norm. All other phenotypes or genotypes are considered variants of this standard, meaning that they deviate from the wild type. The variant may be recessive or dominant to the wild-type allele.

An example of multiple alleles is coat color in rabbits ( [link] ). Here, four alleles exist for the c gene. The wild-type version, C + C + , is expressed as brown fur. The chinchilla phenotype, c ch c ch , is expressed as black-tipped white fur. The Himalayan phenotype, c h c h , has black fur on the extremities and white fur elsewhere. Finally, the albino, or “colorless” phenotype, cc , is expressed as white fur. In cases of multiple alleles, dominance hierarchies can exist. In this case, the wild-type allele is dominant over all the others, chinchilla is incompletely dominant over Himalayan and albino, and Himalayan is dominant over albino. This hierarchy, or allelic series, was revealed by observing the phenotypes of each possible heterozygote offspring.

This illustration shows the four different variants for coat color in rabbits at the c allele. The genotype CC produces the wild type phenotype, which is brown. The genotype c^{ch}c^{ch} produces the chinchilla phenotype, which is black-tipped white fur. The genotype c^{h}c^{h} produces the Himalayan phenotype, which is white on the body and black on the extremities. The genotype cc produces the recessive phenotype, which is white
Four different alleles exist for the rabbit coat color ( C ) gene.

The complete dominance of a wild-type phenotype over all other mutants often occurs as an effect of “dosage” of a specific gene product, such that the wild-type allele supplies the correct amount of gene product whereas the mutant alleles cannot. For the allelic series in rabbits, the wild-type allele may supply a given dosage of fur pigment, whereas the mutants supply a lesser dosage or none at all. Interestingly, the Himalayan phenotype is the result of an allele that produces a temperature-sensitive gene product that only produces pigment in the cooler extremities of the rabbit’s body.

Alternatively, one mutant allele can be dominant over all other phenotypes, including the wild type. This may occur when the mutant allele somehow interferes with the genetic message so that even a heterozygote with one wild-type allele copy expresses the mutant phenotype. One way in which the mutant allele can interfere is by enhancing the function of the wild-type gene product or changing its distribution in the body. One example of this is the Antennapedia mutation in Drosophila ( [link] ). In this case, the mutant allele expands the distribution of the gene product, and as a result, the Antennapedia heterozygote develops legs on its head where its antennae should be.

Questions & Answers

what is nutrition
Joseph Reply
the process of providing or obtaining the food necessary for health and growth.
fatimah
what are micro-organism
Bukiwe Reply
what is immunity
Moriba Reply
What is cross pollination
Moriba
what is pollution
Georgiana Reply
what is pollution
Georgiana
list four type of biological association
Georgiana
Pollution is the release of harmful substances in the atmosphere.
Immanuel
Four biological associations are 1) commensalism 2)symbiosis 3)mutualism 4) parasitism
Immanuel
What is the function of the hormones
Immanuel Reply
What is the other name of filament of a flower
Immanuel
oxygen
Georgiana
list three bone disorders and explain them
Blessings Reply
oxygen
Georgiana
what is self pollination?
Kelvin Reply
well, self-pollination occurs in a single flower which has the organs of both genders. the pollen grains from the anther shall be captured by the stigma of the flower, when released. and the pollen grain shall start grow pollen tube which grows down in the style, to the ovary, ovules and the
Izzati
insides of the ovules. so pretty much the pollen grains being pollinated from the anther to the stigma of the same flower
Izzati
I think
Izzati
the pollen grains being pollinated from the anther to the stigma of the same flower
Izzati
what is animal science
Rose Reply
Animal science is described as "studying the biology of animals that are under the control of humankind".
AMIIRA
don't you guys have Google
Izzati
what is self pollinating
JOSHUA Reply
what is cleistogamy
JOSHUA
Self pollination occurs when the pollen from the anther is deposited on the stigma of the same flower, or another flower on the same plant.
Franklina
Cleistogamy is a type of automatic self pollination of certain plants that can propagate by using non opening, self pollinating flowers
Franklina
Self pollination is the transfer of pollen grain from the anther to the stigma
Immanuel
who is the father of biology
Sean Reply
Aristotle
Lubera
Aristotle
estella
yes Aristotle looking at classification
John
Aristotle
JOSHUA
Aristotle
Grace
please my answer
Sean
Aristotle
Rose
Aristotle
Immanuel
what are prokaryotic cells
Kusubira Reply
having cell that lack membrane - bound nuclei.
Lubera
Pronkayotic cells are cells that lacks nuclear membrane
Immanuel
Please each person should suggest ur answers
Franklina Reply
differentiate between autelogical and synecology
Christiana Reply
differentiate between autelogical and synecology
Christiana Reply
answer please
Edmond
Autecology is the study of individual organism or individual species. It is also known as population ecology. Synecology is the study of group of organisms of different species which are associated together as a unit in form of a community.
Daniel
Autecology is the study of the environment in relation to only one species in contrast while synecology is the study of the environment affecting groups of species coexisting in an area.
Franklina
Describe the factors affecting homotasis
Grace Reply
Fluids and electrolytes Energy and nutrition Immune response mediators
Franklina

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Source:  OpenStax, Biology. OpenStax CNX. Feb 29, 2016 Download for free at http://cnx.org/content/col11448/1.10
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