<< Chapter < Page Chapter >> Page >
 Left illustration shows a cross section of a woody stem undergoing primary growth. At the core of the stem is pith. Toward the outside are egg-shaped vascular bundles. Xylem is located toward the inside of the vascular bundle, and phloem is in the middle. Sclerenchyma cap the outside of the bundle. Right illustration shows a cross section of a woody stem undergoing secondary growth. As in primary growth, the core of the stem is pith. Outside the pith is a ring of secondary xylem. Rounded bundles of primary xylem tissue project from this ring into the pith. Outside the secondary xylem is a ring of secondary phloem tissue. The vascular cambium separates the xylem from the phloem. Outside the secondary phloem is the cortex layer. Bundles of primary phloem project outward from the secondary phloem into the cortex. A cork ring surrounds the cortex. The cork is separated from the cortex by a thin cork cambium. The bark of the tree extends from the vascular cambium to the epidermis.
In woody plants, primary growth is followed by secondary growth, which allows the plant stem to increase in thickness or girth. Secondary vascular tissue is added as the plant grows, as well as a cork layer. The bark of a tree extends from the vascular cambium to the epidermis.

Some plant parts, such as stems and roots, continue to grow throughout a plant’s life: a phenomenon called indeterminate growth. Other plant parts, such as leaves and flowers, exhibit determinate growth, which ceases when a plant part reaches a particular size.

Primary growth

Most primary growth occurs at the apices, or tips, of stems and roots. Primary growth is a result of rapidly dividing cells in the apical meristems at the shoot tip and root tip. Subsequent cell elongation also contributes to primary growth. The growth of shoots and roots during primary growth enables plants to continuously seek water (roots) or sunlight (shoots).

The influence of the apical bud on overall plant growth is known as apical dominance, which diminishes the growth of axillary buds that form along the sides of branches and stems. Most coniferous trees exhibit strong apical dominance, thus producing the typical conical Christmas tree shape. If the apical bud is removed, then the axillary buds will start forming lateral branches. Gardeners make use of this fact when they prune plants by cutting off the tops of branches, thus encouraging the axillary buds to grow out, giving the plant a bushy shape.

Watch this BBC Nature video showing how time-lapse photography captures plant growth at high speed.

Secondary growth

The increase in stem thickness that results from secondary growth is due to the activity of the lateral meristems, which are lacking in herbaceous plants. Lateral meristems include the vascular cambium and, in woody plants, the cork cambium (see [link] ). The vascular cambium is located just outside the primary xylem and to the interior of the primary phloem. The cells of the vascular cambium divide and form secondary xylem (tracheids and vessel elements) to the inside, and secondary phloem (sieve elements and companion cells) to the outside. The thickening of the stem that occurs in secondary growth is due to the formation of secondary phloem and secondary xylem by the vascular cambium, plus the action of cork cambium, which forms the tough outermost layer of the stem. The cells of the secondary xylem contain lignin, which provides hardiness and strength.

In woody plants, cork cambium is the outermost lateral meristem. It produces cork cells (bark) containing a waxy substance known as suberin that can repel water. The bark protects the plant against physical damage and helps reduce water loss. The cork cambium also produces a layer of cells known as phelloderm, which grows inward from the cambium. The cork cambium, cork cells, and phelloderm are collectively termed the periderm    . The periderm substitutes for the epidermis in mature plants. In some plants, the periderm has many openings, known as lenticels , which allow the interior cells to exchange gases with the outside atmosphere ( [link] ). This supplies oxygen to the living and metabolically active cells of the cortex, xylem and phloem.

Questions & Answers

what is nutrients
Patrick Reply
what are the types of skeleton
Tanyi Reply
endo skeleton and exo skeleton
Faith
or types of skeletons There is exo skeleton found in class insecta Exo skeleton found in phylum chordata and lastly hydrostatic skeleton found mainly in earth worms
Daniel
and hydrostatic skeleton
Godbless
what is life?
Jerson Reply
life is when any living thing breath or respirate.
Lucky
I want to join
Patrick
me too
Favour
me two
Monday
me too
Bengazy
me too
Maggie
I here
Yahaya
Does anybody knows another biology app that brings good understand
Lucky
give an example of a phylum where all members have open circulatory
Omoro Reply
anthropoda
Daniel
phylum Anthropoda
Daniel
how do u change the unit of temperature from degree celsius to degree Fahrenheit
Maggie
what are beneficial effects in industrial
Nnamani Reply
create employment
ibrahim
provide food
ibrahim
pollution
ibrahim
dirty the environment
ibrahim
pollution and lack of knowledge about how to manage waste
Alex
what is gametophyte
Ibrahim Reply
what is biology
Basiru Reply
Biology deals with the study of living organisms and their interactions with each other and the environment.
Quartey
biology simply means the study of living and non-livingthings.
Shamstar
what is the meaning of ubiquitous
Hamziyatu Reply
Under what topic is this question
fred
three branches of biology
Alexander Reply
what is the cell
Olajide Reply
what is life
Olajide
cell is the functional and structural unit of life
Shweta
life is a problem solving entity
Atanga
what is the characteristics of plant cell
Quaku
Describe the process that results in the formation of a tetrad.
Michael Reply
what is biology
Ndukwe Reply
biology is the study of life
Oghenemine
what is biology
Jimmy Reply
the study of life
Hamziyatu
what is the Biosphere?
Kabel Reply
I want to know what classifications of living organisms and their names
Quaku
are classified into plants and animals eukaryotic and prokaryotic cell.
Lokwe
what is biology?
Turabi Reply
what is the meaning of Biologists
Turabi
what is the meaning of Biologists
Turabi
what is the meaning of Biologists
Turabi
are those who are specialized in Biology
Nicode
Is the study of living things,
Kabel
what is the ecosystem?
Kabel
bio mean life logus means study so the study of living things or living organisms is called biology
Faraq
Ecosystem consists of the living organisms in a particular area together with the abiotic factors.
Quartey

Get the best Biology course in your pocket!





Source:  OpenStax, Biology. OpenStax CNX. Feb 29, 2016 Download for free at http://cnx.org/content/col11448/1.10
Google Play and the Google Play logo are trademarks of Google Inc.

Notification Switch

Would you like to follow the 'Biology' conversation and receive update notifications?

Ask