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Mitochondria

Eukaryotic cells may contain anywhere from one to several thousand mitochondria, depending on the cell’s level of energy consumption. Each mitochondrion measures 1 to 10 micrometers in length and exists in the cell as a moving, fusing, and dividing oblong spheroid ( [link] ). However, mitochondria cannot survive outside the cell. As the atmosphere was oxygenated by photosynthesis, and as successful aerobic prokaryotes evolved, evidence suggests that an ancestral cell engulfed and kept alive a free-living, aerobic prokaryote. This gave the host cell the ability to use oxygen to release energy stored in nutrients. Several lines of evidence support that mitochondria are derived from this endosymbiotic event. Mitochondria are shaped like a specific group of bacteria and are surrounded by two membranes, which would result when one membrane-bound organism was engulfed by another membrane-bound organism. The mitochondrial inner membrane involves substantial infoldings or cristae that resemble the textured outer surface of certain bacteria.

The transmission electron micrograph shows two round, membrane-bound organelles inside a cell. The organelles are about 400 nm across and have membranes running through the middle of them.
In this transmission electron micrograph of mitochondria in a mammalian lung cell, the cristae, infoldings of the mitochondrial inner membrane, can be seen in cross-section. (credit: modification of work by Louisa Howard; scale-bar data from Matt Russell)

Mitochondria divide on their own by a process that resembles binary fission in prokaryotes. Mitochondria have their own circular DNA chromosome that carries genes similar to those expressed by bacteria. Mitochondria also have special ribosomes and transfer RNAs that resemble these components in prokaryotes. These features all support that mitochondria were once free-living prokaryotes.

Chloroplasts

Chloroplasts are one type of plastid    , a group of related organelles in plant cells that are involved in the storage of starches, fats, proteins, and pigments. Chloroplasts contain the green pigment chlorophyll and play a role in photosynthesis. Genetic and morphological studies suggest that plastids evolved from the endosymbiosis of an ancestral cell that engulfed a photosynthetic cyanobacterium. Plastids are similar in size and shape to cyanobacteria and are enveloped by two or more membranes, corresponding to the inner and outer membranes of cyanobacteria. Like mitochondria, plastids also contain circular genomes and divide by a process reminiscent of prokaryotic cell division. The chloroplasts of red and green algae exhibit DNA sequences that are closely related to photosynthetic cyanobacteria, suggesting that red and green algae are direct descendants of this endosymbiotic event.

Mitochondria likely evolved before plastids because all eukaryotes have either functional mitochondria or mitochondria-like organelles. In contrast, plastids are only found in a subset of eukaryotes, such as terrestrial plants and algae. One hypothesis of the evolutionary steps leading to the first eukaryote is summarized in [link] .

The illustration shows steps that, according to the endosymbiotic theory, gave rise to eukaryotic organisms. In step 1, infoldings in the plasma membrane of an ancestral prokaryote gave rise to endomembrane components, including a nucleus and endoplasmic reticulum. In step 2, the first endosymbiotic event occurred: The ancestral eukaryote consumed aerobic bacteria that evolved into mitochondria. In a second endosymbiotic event, the early eukaryote consumed photosynthetic bacteria that evolved into chloroplasts.
The first eukaryote may have originated from an ancestral prokaryote that had undergone membrane proliferation, compartmentalization of cellular function (into a nucleus, lysosomes, and an endoplasmic reticulum), and the establishment of endosymbiotic relationships with an aerobic prokaryote and, in some cases, a photosynthetic prokaryote to form mitochondria and chloroplasts, respectively.

The exact steps leading to the first eukaryotic cell can only be hypothesized, and some controversy exists regarding which events actually took place and in what order. Spirochete bacteria have been hypothesized to have given rise to microtubules, and a flagellated prokaryote may have contributed the raw materials for eukaryotic flagella and cilia. Other scientists suggest that membrane proliferation and compartmentalization, not endosymbiotic events, led to the development of mitochondria and plastids. However, the vast majority of studies support the endosymbiotic hypothesis of eukaryotic evolution.

The early eukaryotes were unicellular like most protists are today, but as eukaryotes became more complex, the evolution of multicellularity allowed cells to remain small while still exhibiting specialized functions. The ancestors of today’s multicellular eukaryotes are thought to have evolved about 1.5 billion years ago.

Section summary

The first eukaryotes evolved from ancestral prokaryotes by a process that involved membrane proliferation, the loss of a cell wall, the evolution of a cytoskeleton, and the acquisition and evolution of organelles. Nuclear eukaryotic genes appear to have had an origin in the Archaea, whereas the energy machinery of eukaryotic cells appears to be bacterial in origin. The mitochondria and plastids originated from endosymbiotic events when ancestral cells engulfed an aerobic bacterium (in the case of mitochondria) and a photosynthetic bacterium (in the case of chloroplasts). The evolution of mitochondria likely preceded the evolution of chloroplasts. There is evidence of secondary endosymbiotic events in which plastids appear to be the result of endosymbiosis after a previous endosymbiotic event.

Questions & Answers

why are the laws of thermodynamics considered laws of nature and not scientific theories
Yolonda Reply
archea were given their own separate domain because they are?
Yolonda
which of the following is a basic component of all of the others?
Yolonda
which of the following organization levels is the least inclusive
Yolonda
which cell feature is absent in bacterial cells
Yolonda
which metric movies the base unit of measurement by one thousandth (0.001)?
Yolonda
What is biology?
Blessing Reply
List the branches of biology
Blessing
List the branches of biology
Blessing
List the branches of biology
Blessing
List the branches of biology
Blessing
must all prokaryotic cells posses a cell wall?
chris Reply
what is biology?
Cathy Reply
biology is basically the study of life
Robert
that's true
Tonia
it's the study of living things
Tonia
What is a celiac disease
Falase Reply
distinguish between properties and characteristics
Elee Reply
what are organelles
Elee
organelles are substances that makes up a cell
chris
what is biology
Prevail Reply
biology is science that studies life
Elee
it's the study of living and non living things
now that we have an estimate for the diameter of the cell.what estimate can we make about the volume of the cell?
faxhood Reply
why too much insulin result in low blood sugar
Leri Reply
For example, too much exercise can cost you to lose to much weight. Too much insulin will pull to much sugar out of your systemic system into your cells.
Eric
difference between DNA and RNA
Eyitayo Reply
DNA is deoxyribonuclaic acid. Deoxy refers to a lack of oxygen. The Ribose moity is missing an OH group. I think it is missing from the second C of the ring. RNA is ribonucleic acid. DNA has our genetic code in on it. RNA is translated from DNA and carries the blue print for protein synthesis.
Eric
The OH group on RNA prevents it from being reactive. But it is very unstable though. Would you want such a power tool floating around in your body, no. And you have three types of RNA: mRNA, tRNA and rRNA. Please let me know it this helped?😄
Eric
o
Browse
Eric answered this question perfectly
what determines the aeration level in the soil
Shola Reply
what is homeostasis?
Sarita Reply
It means balance in a biological system.
Eric
What is biology
Don Reply
Biology z the study of life
GOLDEN
what's biology
buran
biology is the study of living nd none living organism
Chinaza
Biology is the study of life
Quadri
yes Sir
Said
what's cell biology
Prince
biology is the study of life
Raheal
Biology is a science subject that deals with the study of living things and how they interact with there environment
chris
what is asexual reproduction,?
Awoi Reply
A type of reproduction which does not involve the fusion of gametes or a change in the number of chromosomes
Serena
Reproduction without sex... In which form a single organism or cell makes a copy of itself.
Serena
Please explain the concept of mitosis and meiosis
Serena
I guess you could use it for study buddies and brushing up on what you need to
Serena
what is mitosis
Klp
Asexual reproduction?
Serena
why pepsin and trypsin released in active form?
Yacqub
mitosis is the type cell division in which two daughter cells have same no. of chormosomes
syed
Hi
Don
chromosome number remains the same in mitosis
mc
Ello
Henry
Hii
Nikky
Yrr help me.
Nikky
Physical chemistry..... Koi h jo mujhe physical chem ki notes send kr ske
Nikky
what is asexual reproduction
Targbe
what makes golgi body in plants
Abdulkareem Reply
name the membrane of the plants
Abdulkareem
how can turners syndrome be corrected before birth
Balinda
which animal survive from being preyed just because of being humble, slow, and not aggressive
Balinda
Plants have golgi body's also. Plants are eukaryotic cells. And membrane bound organelles are a characteristic of eukaryotic cells. Moreover golgi body's are creatted from the ER. Also do not forget plants have plastids and animal cells do not.
Eric

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Source:  OpenStax, Concepts of biology. OpenStax CNX. Feb 29, 2016 Download for free at http://cnx.org/content/col11487/1.9
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