<< Chapter < Page Chapter >> Page >
Chlamydia, Spirochetes, Cyanobacteria, and Gram-positive bacteria are described in this table. Note that bacterial shape is not phylum-dependent; bacteria within a phylum may be cocci, rod-shaped, or spiral. (credit “Chlamydia trachomatis”: modification of work by Dr. Lance Liotta Laboratory, NCI; credit “Treponema pallidum”: modification of work by Dr. David Cox, CDC; credit “Phormidium”: modification of work by USGS; credit “Clostridium difficile”: modification of work by Lois S. Wiggs, CDC; scale-bar data from Matt Russell)
Characteristics of the four phyla of archaea are described. Euryarchaeotes includes methanogens, which produce methane as a metabolic waste product, and halobacteria, which live in an extreme saline environment. Methanogens cause flatulence in humans and other animals. Halobacteria can grow in large blooms that appear reddish, due to the presence of bacterirhodopsin in the membrane. Bacteriorhodopsin is related to the retinal pigment rhodopsin. Micrograph shows rod-shaped Halobacterium. Members of the ubiquitous Crenarchaeotes phylum play an important role in the fixation of carbon. Many members of this group are sulfur-dependent extremophiles. Some are thermophilic or hyperthermophilic. Micrograph shows cocci-shaped Sulfolobus, a genus which grows in volcanic springs at temperatures between 75° and 80°C and at a pH between 2 and 3. The phylum Nanoarchaeotes currently contains only one species, Nanoarchaeum equitans, which has been isolated from the bottom of the Atlantic Ocean, and from the a hydrothermal vent at Yellowstone National Park. It is an obligate symbiont with Ignococcus, another species of archaebacteria. Micrograph shows two small, round N. equitans cells attached to a larger Ignococcus cell. Korarchaeotes are considered to be one of the most primitive forms of life and so far have only been found in the Obsidian Pool, a hot spring at Yellowstone National Park. Micrograph shows a variety of specimens from this group which vary in shape.
Archaea are separated into four phyla: the Korarchaeota, Euryarchaeota, Crenarchaeota, and Nanoarchaeota. (credit “Halobacterium”: modification of work by NASA; credit “Nanoarchaeotum equitans”: modification of work by Karl O. Stetter; credit “korarchaeota”: modification of work by Office of Science of the U.S. Dept. of Energy; scale-bar data from Matt Russell)

The plasma membrane

The plasma membrane is a thin lipid bilayer (6 to 8 nanometers) that completely surrounds the cell and separates the inside from the outside. Its selectively permeable nature keeps ions, proteins, and other molecules within the cell and prevents them from diffusing into the extracellular environment, while other molecules may move through the membrane. Recall that the general structure of a cell membrane is a phospholipid bilayer composed of two layers of lipid molecules. In archaeal cell membranes, isoprene (phytanyl) chains linked to glycerol replace the fatty acids linked to glycerol in bacterial membranes. Some archaeal membranes are lipid monolayers instead of bilayers ( [link] ).

This illustration compares phospholipids from Bacteria and Eukarya to those from Archaea. In Bacteria and Eukarya, fatty acids are attached to glycerol by an ester linkage, while in Archaea, isoprene chains are linked to glycerol by an ether linkage. In the ester linkage, the first carbon in the fatty acid chain has an oxygen double-bonded to it, whereas in the ether linkage, it does not. In Archaea, the isoprene chains have methyl groups branching off from them, whereas such branches are absent in Bacteria and Eukarya.  Both types of phospholipids result in similar lipid bilayers.
Archaeal phospholipids differ from those found in Bacteria and Eukarya in two ways. First, they have branched phytanyl sidechains instead of linear ones. Second, an ether bond instead of an ester bond connects the lipid to the glycerol.

The cell wall

The cytoplasm of prokaryotic cells has a high concentration of dissolved solutes. Therefore, the osmotic pressure within the cell is relatively high. The cell wall is a protective layer that surrounds some cells and gives them shape and rigidity. It is located outside the cell membrane and prevents osmotic lysis (bursting due to increasing volume). The chemical composition of the cell walls varies between archaea and bacteria, and also varies between bacterial species.

Bacterial cell walls contain peptidoglycan    , composed of polysaccharide chains that are cross-linked by unusual peptides containing both L- and D-amino acids including D-glutamic acid and D-alanine. Proteins normally have only L-amino acids; as a consequence, many of our antibiotics work by mimicking D-amino acids and therefore have specific effects on bacterial cell wall development. There are more than 100 different forms of peptidoglycan. S-layer    (surface layer) proteins are also present on the outside of cell walls of both archaea and bacteria.

Bacteria are divided into two major groups: Gram positive    and Gram negative    , based on their reaction to Gram staining. Note that all Gram-positive bacteria belong to one phylum; bacteria in the other phyla (Proteobacteria, Chlamydias, Spirochetes, Cyanobacteria, and others) are Gram-negative. The Gram staining method is named after its inventor, Danish scientist Hans Christian Gram (1853–1938). The different bacterial responses to the staining procedure are ultimately due to cell wall structure. Gram-positive organisms typically lack the outer membrane found in Gram-negative organisms ( [link] ). Up to 90 percent of the cell wall in Gram-positive bacteria is composed of peptidoglycan, and most of the rest is composed of acidic substances called teichoic acids . Teichoic acids may be covalently linked to lipids in the plasma membrane to form lipoteichoic acids. Lipoteichoic acids anchor the cell wall to the cell membrane. Gram-negative bacteria have a relatively thin cell wall composed of a few layers of peptidoglycan (only 10 percent of the total cell wall), surrounded by an outer envelope containing lipopolysaccharides (LPS) and lipoproteins. This outer envelope is sometimes referred to as a second lipid bilayer. The chemistry of this outer envelope is very different, however, from that of the typical lipid bilayer that forms plasma membranes.

Questions & Answers

what is the protein found in the blood?
Tobias Reply
globin
Joelia
Globin
what is parasitic movement
Emmanuel Reply
Parasitic movement is a problem for all of us. So is its companion, parasitic tension. Parasitic movement is the excess contraction of muscles that you don't actually need to complete an action.
freya
what are eukaryotic cells
Thiza Reply
what is the mean of pair of chromosomes
Kazula Reply
hi
Lagos
23 haploid and 23diploid
Patson
how are you studying in this quarantine? .. how are you keeping yourselves motivated?
sivajijadhav @815.com
good morning guyz
Joelia
morning
Kazula
hi
Justin
Good
Angela
tell me if you know what can be used...than reading pls hint me pls 🙏🙏🙏
Angela
what is the important of sex
Aremu Reply
why did human being need sex?
Aremu
because he/she have feelings
Chripine
reproduction...to make more
Yazi
due to active harmon
Manish
One important of sex is to reproduce
Emma
to ensure the countinuty of life
Yusuf
all of you are right
Edith
what is momentum
Asiya Reply
The strength or force that allows something to continue or grow stronger or faster as time pass
Emma
What is Centripetal Force?
Justin
centrepital force is the inward force required to keep a body moving with constant speed in a circular path
Yusuf
what is the test for protein
Takii Reply
List four condition necessary for seed germination
Tedeka Reply
water,air(oxygen),light,temperature
Hassan
water, light, oxygen and temperature
lilchris
water, oxygen, light temperature
Mike
water oxygen light and temperature
John
importance of biology
Alabina Reply
importance of boilogy
Alabina
what is soil
Amina Reply
soil is the upper part of the earth
Alabina
what is importance of studying biology
Alabina
soil is the uppermost layer of the earth on which plant grows
Yusuf
soil is defined as the thin surface of the upper most layer of the earth crust on which plants grow
Thanni
soil is the upper part of the earth which plants grow on
Emma
differences between euglenoid and amoeboid
Grace Reply
what are the difference between aerobic and anaerobic respiration?
Maxwell Reply
Aerobic respiration involves the use of oxygen whiles anaerobic respiration does not involve the use of oxygen
Quabena
what is assmilation
Lucy Reply
what is cell
Manish Reply
cell is the structural and functional unit of life or living things
hamid
where anaerobic respiration occurre?
Manish
in cell
Manish
in cells?
Manish
where anaerobic respiration occurre in cell?
Manish

Get the best Biology course in your pocket!





Source:  OpenStax, Biology. OpenStax CNX. Feb 29, 2016 Download for free at http://cnx.org/content/col11448/1.10
Google Play and the Google Play logo are trademarks of Google Inc.

Notification Switch

Would you like to follow the 'Biology' conversation and receive update notifications?

Ask