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Photo shows dolphins lying on the edge of their tank, being fed fish by their trainers.
The training of dolphins by rewarding them with food is an example of positive reinforcement operant conditioning. (credit: Roland Tanglao)

Cognitive learning

Classical and operant conditioning are inefficient ways for humans and other intelligent animals to learn. Some primates, including humans, are able to learn by imitating the behavior of others and by taking instructions. The development of complex language by humans has made cognitive learning    , the manipulation of information using the mind, the most prominent method of human learning. In fact, that is how students are learning right now by reading this book. As students read, they can make mental images of objects or organisms and imagine changes to them, or behaviors by them, and anticipate the consequences. In addition to visual processing, cognitive learning is also enhanced by remembering past experiences, touching physical objects, hearing sounds, tasting food, and a variety of other sensory-based inputs. Cognitive learning is so powerful that it can be used to understand conditioning in detail. In the reverse scenario, conditioning cannot help someone learn about cognition.

Classic work on cognitive learning was done by Wolfgang Köhler with chimpanzees. He demonstrated that these animals were capable of abstract thought by showing that they could learn how to solve a puzzle. When a banana was hung in their cage too high for them to reach, and several boxes were placed randomly on the floor, some of the chimps were able to stack the boxes one on top of the other, climb on top of them, and get the banana. This implies that they could visualize the result of stacking the boxes even before they had performed the action. This type of learning is much more powerful and versatile than conditioning.

Cognitive learning is not limited to primates, although they are the most efficient in using it. Maze running experiments done with rats by H.C. Blodgett in the 1920s were the first to show cognitive skills in a simple mammal. The motivation for the animals to work their way through the maze was a piece of food at its end. In these studies, the animals in Group I were run in one trial per day and had food available to them each day on completion of the run ( [link] ). Group II rats were not fed in the maze for the first six days and then subsequent runs were done with food for several days after. Group III rats had food available on the third day and every day thereafter. The results were that the control rats, Group I, learned quickly, and figured out how to run the maze in seven days. Group III did not learn much during the three days without food, but rapidly caught up to the control group when given the food reward. Group II learned very slowly for the six days with no reward to motivate them, and they did not begin to catch up to the control group until the day food was given, and then it took two days longer to learn the maze.

A diagram shows a rat maze that has several turns and dead ends. Next to this maze is a graph showing the average error of the rats navigating the maze plotted versus the day of the experiment. Rats that are rewarded at the end of each run learn the maze quickly, and the number of errors they make in navigating the maze rapidly drops from six on day one to one on day eleven. Rats that are not rewarded on the first three days but are rewarded after day three learn the maze slowly at first, but quickly after the reward is present. Rats that are not rewarded on the first six days but are rewarded after day six gradually reduce the number of errors over the first three days, but rapidly reduce their errors after an award is present.
Group I (the green solid line) found food at the end of each trial, group II (the blue dashed line) did not find food for the first 6 days, and group III (the red dotted line) did not find food during runs on the first three days. Notice that rats given food earlier learned faster and eventually caught up to the control group. The orange dots on the group II and III lines show the days when food rewards were added to the mazes.

It may not be immediately obvious that this type of learning is different than conditioning. Although one might be tempted to believe that the rats simply learned how to find their way through a conditioned series of right and left turns, E.C. Tolman proved a decade later that the rats were making a representation of the maze in their minds, which he called a “cognitive map.” This was an early demonstration of the power of cognitive learning and how these abilities were not just limited to humans.


Sociobiology is an interdisciplinary science originally popularized by social insect researcher E.O. Wilson in the 1970s. Wilson defined the science as “the extension of population biology and evolutionary theory to social organization.” Edward O. Wilson. On Human Nature (1978; repr., Cambridge: Harvard University Press, 2004), xx. The main thrust of sociobiology is that animal and human behavior, including aggressiveness and other social interactions, can be explained almost solely in terms of genetics and natural selection. This science is controversial; noted scientist such as the late Stephen Jay Gould criticized the approach for ignoring the environmental effects on behavior. This is another example of the “nature versus nurture” debate of the role of genetics versus the role of environment in determining an organism’s characteristics.

Sociobiology also links genes with behaviors and has been associated with “biological determinism,” the belief that all behaviors are hardwired into our genes. No one disputes that certain behaviors can be inherited and that natural selection plays a role retaining them. It is the application of such principles to human behavior that sparks this controversy, which remains active today.

Section summary

Behaviors are responses to stimuli. They can either be instinctual/innate behaviors, which are not influenced by the environment, or learned behaviors, which are influenced by environmental changes. Instinctual behaviors include mating systems and methods of communication. Learned behaviors include imprinting and habituation, conditioning, and, most powerfully, cognitive learning. Although the connection between behavior, genetics, and evolution is well established, the explanation of human behavior as entirely genetic is controversial.

Questions & Answers

there are 3 trimester in human pregnancy
I don't know answer of this question can u help me
what is a cell
Fatima Reply
what is genetic
Janet Reply
I join
what are the branchas of biology
Prisca Reply
zoology, ecology
genetics, microbiology,botany and embryology
what is a cell
Kulunbawi Reply
cell is smallest unit of life. cells are often cell the building blocks of life...
the first twenty element
Orapinega Reply
what are the characteristics of living things?
growth,respiration,nutrition,sensitivity, movement,irritability, excretion,death.
What is the difference between adaptation and competition in animals
Adeyemi Reply
What is biology
it is a natural science stadey about living things
Biology is the bronch of science which deals with the study of life is called biology
what is the x in 300 stands for?
Ogbudu Reply
the properties of life
Clarinda Reply
response to the environment, reproduction, homeostasis, growth,energy processing etc.....
what is reproduction
Reproduction is a fundamental feature of all known life,each individual organism exist as a result of re production.....or else Multiplying...
a complete virus particle known as
Darlington Reply
These are formed from identical protein subunitscalled capsomeres.
fabace family plant name
Pushpam Reply
in eukaryotes ...protein channel name which transport protein ...
Pushpam Reply
in bacteria ...chromosomal dna duplicate structure called
what is a prokaryotic cell and a eukaryotic cell
Matilda Reply
There are two types of cells. Eukaryotic and Prokaryotic cells. Prokaryotic cells don't have a nucleus or membrane enclosed organelles (little organs within that cell). They do however carry genetic material but it's not maintained in the nucleus. Prokaryotic cells are also one celled.
Prokaryotic cells are one celled (single celled).
Prokaryotic cells are Bacteria and Archea
Prokaryotic cells are smaller than Eukaryotic cells.
Eukaryotic cells are more complex. They are much bigger than Prokaryotic cells.
Eukaryotic cells have a nucleus and membrane bound organelles.
Eukaryotic cells are animals cells which also includes us.
Eukaryotic cells are also multicellular.
nice explaination
eukaryotic cells are individual cells .. but eukaryotes are multicellular organisms which consist of many different types of eukaryotic cells
also eukaryotic cells have mitochondria. prokaryotic cells do not
in prokaryotes only ribosomes are present... in eukaryotes mitochondria ...glogi bodies ..epidermis .....prokaryotes one envelop but eukaryotes compartment envelop....envelop mean membrane bound organelles......
prokaryotic cell are cells dat have no true nuclei i.e no cell membrane while eukaryotic cell are cell dat have true nuclei i.e have cell membrane
we have 46 pair of somatic cell and 23 pair of chromosomes in our body, pls can someone explain it to me. pls
Matilda Reply
we have 22 pairs of somatic chromosomes and one pair of sex chromosome
we have 23 pairs of chromosomes,22 pairs of somatic and one pair of sex chromosomes
23 chromosomes from dad & 23 chromosomes from mom 23 +23=46 total chromosomes
X & Y chromosomes are called sex cells, the very presence of a Y chromosome means the person is Male.
XX Female XY Male
If a Karyotype has more than 46 Chromosomes then nondisjunction occured. For example, having an extra chromosome 21 will cause Down Syndrome.
in mammal state the different vertebrae and their location in the body
Igbinigie Reply

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