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Photo a shows Rana aurora, a beige frog with green spots. Photo b shows Rana boylii, a brown frog.
These two related frog species exhibit temporal reproductive isolation. (a) Rana aurora breeds earlier in the year than (b) Rana boylii . (credit a: modification of work by Mark R. Jennings, USFWS; credit b: modification of work by Alessandro Catenazzi)

In some cases, populations of a species move or are moved to a new habitat and take up residence in a place that no longer overlaps with the other populations of the same species. This situation is called habitat isolation    . Reproduction with the parent species ceases, and a new group exists that is now reproductively and genetically independent. For example, a cricket population that was divided after a flood could no longer interact with each other. Over time, the forces of natural selection, mutation, and genetic drift will likely result in the divergence of the two groups ( [link] ).

Illustration A shows the black Gryllus pennsylvanicus cricket on sandy soil, and illustration B shows the beige Gryllus firmus cricket in grass.
Speciation can occur when two populations occupy different habitats. The habitats need not be far apart. The cricket (a) Gryllus pennsylvanicus prefers sandy soil, and the cricket (b) Gryllus firmus prefers loamy soil. The two species can live in close proximity, but because of their different soil preferences, they became genetically isolated.

Behavioral isolation occurs when the presence or absence of a specific behavior prevents reproduction from taking place. For example, male fireflies use specific light patterns to attract females. Various species of fireflies display their lights differently. If a male of one species tried to attract the female of another, she would not recognize the light pattern and would not mate with the male.

Other prezygotic barriers work when differences in their gamete cells (eggs and sperm) prevent fertilization from taking place; this is called a gametic barrier    . Similarly, in some cases closely related organisms try to mate, but their reproductive structures simply do not fit together. For example, damselfly males of different species have differently shaped reproductive organs. If one species tries to mate with the female of another, their body parts simply do not fit together. ( [link] ).

Illustrations show four different types of damselfly reproductive organs. Each organ has a hook, but the shape and length of the hook varies, as does the shape of the organ itself.
The shape of the male reproductive organ varies among male damselfly species, and is only compatible with the female of that species. Reproductive organ incompatibility keeps the species reproductively isolated.

In plants, certain structures aimed to attract one type of pollinator simultaneously prevent a different pollinator from accessing the pollen. The tunnel through which an animal must access nectar can vary widely in length and diameter, which prevents the plant from being cross-pollinated with a different species ( [link] ).

Illustration a shows a bee collecting pollen from a bright purple foxglove flower. The bee’s body fits inside the bell-like flower. Illustration B shows a hummingbird drinking nectar from a long tube-like trumpet creeper flower.
Some flowers have evolved to attract certain pollinators. The (a) wide foxglove flower is adapted for pollination by bees, while the (b) long, tube-shaped trumpet creeper flower is adapted for pollination by humming birds.

When fertilization takes place and a zygote forms, postzygotic barriers can prevent reproduction. Hybrid individuals in many cases cannot form normally in the womb and simply do not survive past the embryonic stages. This is called hybrid inviability because the hybrid organisms simply are not viable. In another postzygotic situation, reproduction leads to the birth and growth of a hybrid that is sterile and unable to reproduce offspring of their own; this is called hybrid sterility.

Habitat influence on speciation

Sympatric speciation may also take place in ways other than polyploidy. For example, consider a species of fish that lives in a lake. As the population grows, competition for food also grows. Under pressure to find food, suppose that a group of these fish had the genetic flexibility to discover and feed off another resource that was unused by the other fish. What if this new food source was found at a different depth of the lake? Over time, those feeding on the second food source would interact more with each other than the other fish; therefore, they would breed together as well. Offspring of these fish would likely behave as their parents: feeding and living in the same area and keeping separate from the original population. If this group of fish continued to remain separate from the first population, eventually sympatric speciation might occur as more genetic differences accumulated between them.

This scenario does play out in nature, as do others that lead to reproductive isolation. One such place is Lake Victoria in Africa, famous for its sympatric speciation of cichlid fish. Researchers have found hundreds of sympatric speciation events in these fish, which have not only happened in great number, but also over a short period of time. [link] shows this type of speciation among a cichlid fish population in Nicaragua. In this locale, two types of cichlids live in the same geographic location but have come to have different morphologies that allow them to eat various food sources.

Illustrations show two species of cichlid fish which are similar in appearance except that one has thin lips, and one has thick lips.
Cichlid fish from Lake Apoyeque, Nicaragua, show evidence of sympatric speciation. Lake Apoyeque, a crater lake, is 1800 years old, but genetic evidence indicates that the lake was populated only 100 years ago by a single population of cichlid fish. Nevertheless, two populations with distinct morphologies and diets now exist in the lake, and scientists believe these populations may be in an early stage of speciation.

Section summary

Speciation occurs along two main pathways: geographic separation (allopatric speciation) and through mechanisms that occur within a shared habitat (sympatric speciation). Both pathways isolate a population reproductively in some form. Mechanisms of reproductive isolation act as barriers between closely related species, enabling them to diverge and exist as genetically independent species. Prezygotic barriers block reproduction prior to formation of a zygote, whereas postzygotic barriers block reproduction after fertilization occurs. For a new species to develop, something must cause a breach in the reproductive barriers. Sympatric speciation can occur through errors in meiosis that form gametes with extra chromosomes (polyploidy). Autopolyploidy occurs within a single species, whereas allopolyploidy occurs between closely related species.

Art connections

[link] Which is most likely to survive, offspring with 2 n +1 chromosomes or offspring with 2 n -1 chromosomes?

[link] Loss of genetic material is almost always lethal, so offspring with 2 n +1 chromosomes are more likely to survive.

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Questions & Answers

who is the father of biology
Sean Reply
Aristotle
Lubera
Aristotle
estella
what are prokaryotic cells
Kusubira Reply
having cell that lack membrane - bound nuclei.
Lubera
Please each person should suggest ur answers
Franklina Reply
differentiate between autelogical and synecology
Christiana Reply
differentiate between autelogical and synecology
Christiana Reply
answer please
Edmond
Autecology is the study of individual organism or individual species. It is also known as population ecology. Synecology is the study of group of organisms of different species which are associated together as a unit in form of a community.
Daniel
Autecology is the study of the environment in relation to only one species in contrast while synecology is the study of the environment affecting groups of species coexisting in an area.
Franklina
Describe the factors affecting homotasis
Grace Reply
Fluids and electrolytes Energy and nutrition Immune response mediators
Franklina
prove starch in a mango leaf
Ruth Reply
prove starch in a leaf
Ruth
draw and name the plant cell
Nalongo
wat is classification
Nalongo
levels of classification
Nalongo
classification of what
Ezenwa
Nidhipal
Nidhi
Iodine solution is used to test leaves for the presence of starch.
Franklina
Identify the caste of termites or Honey bees
Mensah Reply
Soldier Winged reproductive Worker King and queen
Franklina
what is biology
Jackson Reply
biology is a study of living organism
Brilliant
what is the myelin sheath?
Myelin sheath is an insulting layer that forms around nerves, including those in the brain and spinal cord.
Franklina
description on how an enzyme such as pepsin breaks down a substrate
linister Reply
The specific reaction catalyzed by pepsin is the acid hydrolysis of the peptide bond. This reaction will break down protein into smaller units to enable the digestive process
Franklina
what is biology
Kanzy Reply
study of life
Tufail
the study of life..meaning studying all aspects of life in humans,animals and everything inside the global
Nalukui
study of life
Memory
is the branch of science which deal with the study of living and non living things
David
Is the study of living organisms or living things
Franklina
draw the taxonomic hierarchy of classification
KC Reply
Kingdom Class Order Family Genus Species
Franklina
explain the following bush burning, deforestation, over grazing
Thomas Reply
what is biodiversity
Tracy
No idea
nana
Please what are alleles
nana
Deforestation: this is the cutting down of trees without planting new ones
Brilliant
over grazing is the continuously planting of crops in one particular land.
Friday
what is afforestation
Friday
Bush burning is the removal of the natural vegetation cover, that protects the soil surface through the use of fire. Deforestation is the actor process of establishing a forest especially on land not previously forested.
Franklina
Over grazing is the practice of grazing toi many livestock for too long a period on a land unable to recover its vegetation.
Franklina
what is physiology
Joseph Reply
the study of physical things
Nalukui
dealing with those things which you can see by your neck eyes
Odwa
Further explanation please
nana
the study of things tt interact with time,energy
David
Is the scientific study of chemistry and physics of the structure of the body
Franklina

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Source:  OpenStax, Biology. OpenStax CNX. Feb 29, 2016 Download for free at http://cnx.org/content/col11448/1.10
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